1. Laboratory of Biology and Environment, Campus Chaab-Erssas, Biopôle, University Brothers Mentouri-Constantine1, Constantine, Algeria
2. Institute of Nutrition, Food and Agri-Food Technologies, University Brothers Mentouri-Constantine 1, Constantine, Algeria
3. Department of Invertebrate Animals, Natural History Museum of the Burgergemeinde Bern, Bern, Switzerland
4. Institute of Ecology and Evolution, University of Bern, Bern, Switzerland