1. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design, Beijing Forestry University, Beijing, China
2. College of Forestry, Beijing Forestry University, Beijing, China
3. College of Chemistry, Chemical Engineering and Resource Utilization, Northeast Forestry University, Harbin, China
4. State Key Laboratory of Tree Genetics and Breeding, Northeast Forestry University, Harbin, China
5. Forest Biotechnology Group, Department of Forestry and Environmental Resources, North Carolina State University, Raleigh, NC, USA