1. a Department of Plant and Soil Science , University of Aberdeen , Aberdeen , AB9 1FX , UK
2. b Station de Pathologie Végétale, INRA, Ministere de l'Agriculture, Domaine Saint-Maurice , B.P. 94, 84143 , Montfavet , France
3. c Department of Applied Biochemistry and Food Science , University of Nottingham, Sutton Bonington Campus , Loughborough , LE12 5RD , UK