1. Faculty of Rangeland and Watershed Management, Gorgan University of Agricultural Science and Natural Resources, Gorgan, Iran;
2. Department of Range and Watershed Management, Faculty of Natural Resources and Environment, Ferdowsi University of Mashhad, Mashhad, Iran;
3. School of Plant Sciences, The University of Arizona, Tucson, AZ, USA;
4. Desert Studies Faculty, Semnan University, Semnan, Iran