1. Department of Pathology and Cell Biology, Columbia University, New York, NY, USA
2. National Institute on Aging Intramural Research Program, NIH, Baltimore, MD, USA
3. Division of Neuroscience, Oregon National Primate Research Center, Beaverton, OR, USA
4. Department of Ophthalmology, Columbia University, New York, NY, USA
5. Department of Clinical Neuroscience, Karolinska Institutet, Stockholm, Sweden