1. Royal Botanic Garden Edinburgh, Edinburgh, Scotland, UK
2. Botanical Museum & Department of Biosciences, Viikki Plant Science Centre, University of Helsinki, Helsinki, Finland
3. Taiwan Endemic Species Research Institute, JiJi, Nantou County, Taiwan
4. Herbarium, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China