1. Bryophyte Surveys Ltd, 20 Thornleigh Road, Bristol, Bristol BS7 8PH, UK
2. UK Centre for Ecology and Hydrology, Bangor, LL57 2UW, UK
3. Bangor University, Bangor, LL57 2DG, UK
4. Natural Resources Wales, Bangor, LL57 2DW, UK
5. Ty Newydd, Church Street, Newborough, LL61 6SD, UK