1. a NOAA Coastal Services Center , 2234 South Hobson Avenue , Charleston, South Carolina 29405‐2413, USA
2. b George Mason University, Department of Geography , MSN 1E2 , Fairfax, Virginia 22030‐4444, USA
3. c State of Alaska, Division of Wildlife Conservation , 333 Raspberry Road , Anchorage, Alaska 99518, USA