1. a School of Conservation Sciences , Bournemouth University , Talbot Campus, Fern Barrow, Poole, Dorset, BH12 5BB, UK
2. b Division of Geography , Northumbria University , Lipman Building, Newcastle upon Tyne NE1 8ST, UK
3. c Centre for Earth and Environmental Science Research , Kingston University , Penryhn Road, Surrey KT1 2EE, UK
4. d School of Environment and Life Sciences , University of Salford , Salford, Greater Manchester M5 4WT, UK