1. a School of Agricultural Science , University of Tasmania , Private Bag 12, Hobart, 7001 Tasmania
2. b Forest Resources Research , PO Box 100, Beecroft, NSW, 2119, Australia
3. c CRC Forestry , Private Bag 12, Hobart, 7001 Tasmania
4. d Ensis (the joint forces of CSIRO and SCION) , Forest Protection and Biosecurity , Private Bag 12, Hobart, 7001 Tasmania