1. a Department of Forestry , Instituto Superior de Agronomia , Tapada da Ajuda, Lisboa, 1349-017, Portugal E-mail:
2. d Tropical Research Institute , Travessa do Conde da Ribeira 9, Lisboa, 1300-142, Portugal E-mail:
3. e Faculty of Agronomy and Forestry , Universidade Eduardo Mondlane , Campus Universit@rio edificio 1, C.P.257, Maputo, Mozambique E-mail:
4. f Servi@os Provinciais de Florestas e Fauna Bravia , C.P.36 Nampula, Mozambique