1. a Quantitative Fisheries Center, Department of Fisheries and Wildlife , Michigan State University , East Lansing , Michigan , 48824 , USA
2. b Chesapeake Biological Laboratory , University of Maryland Center for Environmental Science , Solomons , Maryland , 20688 , USA
3. c Department of Fisheries and Wildlife , USA
4. d Quantitative Fisheries Center, Michigan State University , East Lansing , Michigan , 48824 , USA