1. Department of Plant Genetics and Breeding, Faculty of Agriculture, Tarbiat Modares University, Tehran, Iran
2. Research Faculty of Agriculture, Hokkaido University, Sapporo Japan
3. Department of Agronomy, Faculty of Agriculture, Tarbiat Modares University, Tehran, Iran
4. Department of Cell and Metabolic Biology, Leibniz Institute of Plant Biochemistry, Friedrich-Schiller- University, Jena, Germany
5. Department of Plant Pathology, Faculty of Agriculture, Tarbiat Modares University, Tehran, Iran