1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, People’s Republic of China
2. Department of Zoology, Oxford University, Oxford, OX1 3PS, UK
3. Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, KS, USA
4. Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA