1. Research and Education Department, Gunung Mulu National Park, Mulu, Malaysia
2. Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, Kota Samarahan, Malaysia
3. Department of Life Science, National Taiwan Normal University, Taipei, Taiwan
4. BiBio Research Group, Natural Sciences Museum of Granollers, Granollers, Spain
5. Department of Biology, University of Oxford, Oxford, UK