1. Biodiversity Research & Assessment, Ezemvelo KwaZulu-Natal Wildlife, Montrose, South Africa
2. Centre for Functional Biodiversity, School of Life Sciences, University of KwaZulu-Natal, Scottsville, South Africa
3. African Ecological Conservation Projects, Pretoria, South Africa
4. Department of Nature Conservation, Tshwane University of Technology, Pretoria, South Africa.