1. Unit for Environmental Sciences and Management, North-West University, Potchefstroom, South Africa
2. South African Institute for Aquatic Biodiversity, Makhanda, South Africa
3. Department of Zoology and Entomology, Free State University, Bloemfontein, 9301, South Africa
4. Museum für Naturkunde Berlin - Leibniz Institute for Evolution and Biodiversity Science, Berlin, Germany.