1. a Department of Environmental Science (Biogeography) , University of Basel , Basel , Switzerland
2. b Tropical Biodiversity Section , Museo Tridentino di Scienze Naturali , Trento , Italy
3. c Department of Zoology and Marine Biology , University of Dar es Salaam , Dar es Salaam , Tanzania
4. d Institut für Spezielle Zoologie und Evolutionsbiologie mit Phyletischem Museum , Friedrich Schiller Universität Jena , Jena , Germany