1. Department of Zoology, University of Johannesburg, Johannesburg, South Africa
2. South African National Biodiversity Institute, Kirstenbosch Research Centre, Cape Town, South Africa
3. Department of Biodiversity and Conservation Biology, University of the Western Cape, Cape Town, South Africa
4. Department of Animal and Plant Systematics, National Museum, Bloemfontein, South Africa
5. Department of Zoology and Entomology, University of the Free State, Bloemfontein, South Africa