1. National Zoological Garden, South African National Biodiversity Institute, PO Box 754, Pretoria, 0001, South Africa
2. Genetics Department, University of the Free State, PO Box 339, Bloemfontein, 9300 South Africa
3. Department of Biology, University of New Mexico, Albuquerque, NM, 87131-0001, USA
4. Threatened Amphibian Programme, Endangered Wildlife Trust, Building K2, Pinelands Office Park, Ardeer Road, Modderfontein, 1609, South Africa
5. Unit for Environmental Sciences and Management, North-West University, Potchefstroom, 2520, South Africa
6. South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown 6140
7. Department of Zoology, University of Venda, Thohoyandou, South Africa