1. Division of Environmental Forest Science, Gyeongsang National University, Jinju, South Korea;
2. Department of Seed and Seedling Management, National Forest Seed Variety Center, Chungju, South Korea;
3. Plant Conservation Division, Korea National Arboretum of the Korea Forest Service, Pocheon, South Korea;
4. Institute of Agriculture and Life Science, Gyeongsang National University, Jinju, South Korea