1. Graduate School of Agriculture, Kindai University, Nara, Japan;
2. Nara Prefecture Agricultural Research and Development Center, Nara, Japan
3. School of Environmental Science, The University of Shiga Prefecture, Hikone, Japan
4. Faculty of Agriculture, Ryukoku University, Shiga, Japan
5. National Agricultural Research Center for Hokkaido Region, Upland Agriculture Research Center, Hokkaido, Japan