1. Key Laboratory of Economic Plants and Biotechnology, and Yunnan Key Laboratory for Wild Plant Resources, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
2. Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin 05282, Myanmar
3. State Key Laboratory of Phytochemistry and Plant Resources in West China, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
4. College of Life and Environmental Sciences, Minzu University of China, Beijing 100081, China