1. a Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou , 510650 , China
2. b Guangdong Provincial Key Lab of Biotechnology for Plant Development, South China Normal University , Guangzhou , 510631 , China
3. c College of Chemical and Environmental Engineering, Chongqing Three Gorges University , Chongqing , 404000 , China
4. d Laboratory of NMR Analysis and Measurement, South China Sea Institute of Oceanology, Chinese Academy of Sciences , Guangzhou , 510301 , China