1. a Department of Marine Biotechnology and Resources , National Sun Yat-sen University , Kaohsiung, 804, Taiwan
2. b Department of Microbiology , Kaohsiung Medical University , Kaohsiung, 807, Taiwan
3. c Institute of Oceanography, National Taiwan University , Taipei, Taiwan
4. d Asia-Pacific Ocean Research Center, National Sun Yat-sen University , Kaohsiung, Taiwan