1. School of Medical Sciences, Örebro University, Örebro, Sweden
2. School of Science and Technology, Örebro University, Örebro, Sweden
3. Department of Anatomy, Physiology and Cell Biology, School of Veterinary Medicine, University of California Davis, Davis, CA, USA
4. Turku Bioscience Centre, University of Turku and Åbo Akademi University, Turku, Finland
5. Department of Gastroenterology, Faculty of Medicine and Health, Örebro University, Örebro, Sweden