1. Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, PA, USA
2. Department of Medicine, Division of Experimental Medicine, University of California, San Francisco, CA, USA
3. Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, PA, USA
4. Department of Ophthalmology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA
5. Department of Pathology, University of California, San Francisco, CA, USA
6. Department of Otorhinolaryngology—Head and Neck Surgery, Perelman School of Medicine at The University of Pennsylvania, Philadelphia, PA, USA
7. The Corporal Michael J. Crescenz VA Medical Center Surgical Service, Philadelphia, PA, USA
8. Monell Chemical Senses Center, Philadelphia, PA, USA