1. Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Menglun, Mengla, Yunnan, 666303, China
2. Faculty of Resource Science & Technology, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia
3. Harvard University Herbaria, 22 Divinity Avenue, Cambridge, MA 02138, USA
4. Ludwig-Maximilians-Universität München, Department Biologie I, Systematische Botanik und Mykologie, Menzinger Straße 67, 80638 München, Germany
5. Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, 88400 Kota Kinabalu, Sabah, Malaysia