1. Department of Ophthalmology and Visual Sciences, University of Iowa, Iowa City, Iowa, USA
2. Molecular Otolaryngology and Renal Research Laboratories, Department of Otolaryngology, Head and Neck Surgery, University of Iowa Hospitals and Clinics, Iowa City, Iowa, USA
3. Department of Molecular Physiology & Biophysics, University of Iowa College of Medicine, Iowa City, Iowa, USA
4. Department of Ophthalmology and Visual Sciences, University of British Columbia, Vancouver, British Columbia, Canada
5. Department of Neurology, University of Iowa, Iowa City, Iowa, USA
6. The University of Iowa Institute for Vision Research, Iowa City, Iowa, USA