1. Miyazu Field Station, Japan Fisheries Research and Education Agency, Miyazu, Kyoto, Japan
2. Fishing Vessel Taikimaru, Echizen Town Fisheries Cooperative Association, Echizen, Fukui, Japan
3. Niigata Field Station, Japan Fisheries Research and Education Agency, Niigata, Japan
4. Department of Marine Biosciences, Tokyo University of Marine Science and Technology, Tokyo, Japan