1. A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch, Russian Academy of Sciences, Vladivostok, Russia
2. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok, Russia
3. School of Natural Sciences, Far Eastern Federal University, Vladivostok, Russia
4. Department of Biology, St. Francis Xavier University, Antigonish, NS, Canada