1. Department of Plant and Animal Biology, Faculty of Biological Science and Technology, University of Isfahan, Isfahan, Iran
2. Botany Research Division, Research Institute of Forests and Rangelands, Agricultural Research, Education and Extension Organization (AREEO), Tehran, Iran
3. Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria
4. Leibniz Institute of Plant Genetics and Crop Research (IPK), Gatersleben, Germany
5. German Centre of Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, Germany