1. Laboratory of Extremophile Plants, Centre of Biotechnology of Borj Cedria (CBBC), Hammam-Lif, Tunisia
2. Faculty of Sciences, Department of Life Sciences, University of Gabes, Gabes, Tunisia
3. Research Unit Biodiversity and Valorization of Bioresources in Arid Zones, Faculty of Sciences of Gabes, University of Gabes, Gabes, Tunisia