1. Department of Genetics and Biotechnology, University of Trás-os-Montes e Alto Douro, 5000-801 Vila Real, Portugal;
2. Centre for the Research and Technology of Agro-Environmental and Biological Sciences, University of Trás-os-Montes e Alto Douro, 5000-801 Vila Real, Portugal;
3. Herbarium, Department of Biology and Environment, University of Trás-os-Montes e Alto Douro, 5000-801 Vila Real, Portugal;
4. Evolution, Taxonomy and Conservation Group (ECOMED), Department of Botany, University of Salamanca, 37007 Salamanca, Spain