1. Faculty of Arts and Sciences, Department of Biology, Zonguldak Bülent Ecevit University, Zonguldak, Turkey
2. Laboratory of Lichenology and Bryology, Komarov Botanical Institute of the Russian Academy of Sciences, Saint Petersburg, Russia
3. Faculty of Forestry, Department of ForestEngineering, Çankırı Karatekin University, Çankırı, Turkey