1. Klinkhardt, M. and Tesche, M. and Greven, H.(1995) Database of fish chromosomes. In Westarp Wissenschaften. ( pp. 273 ). Magdeburg
2. Hensley, D. (1997) An overview of the systematics and biogeography of the flatfishes J. Sea Res., 37, pp. 187 - 194.
3. Fukuoka, H. and Niiyama, H. (1970) Notes on the somatic chromosomes of ten species of pleuronetid fishes Chromosome Inf Serv., 11, pp. 18 - 19.
4. LeGrande, W. (1975) Karyology of six species of Louisiana flatfishes (Pleuronectiformes, Osteichthyes) Copeia, 1975(3), pp. 516 - 522.
5. Sakamoto, K. and Nishikawa, S. (1980) Choromosome of three flatfishers (Pleuronectiformes) Japan. J. Ichthy., 27, pp. 268 - 272.