1. NSW National Parks and Wildlife Service, Office of Environment and Heritage, Department of Premier and Cabinet, Hurstville, NSW 2220, Australia
2. Centre for Ecosystem Science, University of New South Wales, Sydney, Australia
3. Research Institute for Environment and Livelihoods, Charles Darwin University, Darwin, Australia
4. School of Biological Sciences, University of Aberdeen, Aberdeen, United Kingdom
5. Department of Environmental Sciences, Macquarie University, Sydney, Australia