1. School of Science, Mae Fah Luang University, Chiang Rai, Thailand
2. Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, People’s Republic of China
3. School of Biology and Chemistry, Pu'er University, Pu’er 665000, China
4. Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand
5. Center of Excellence in Biotechnology Research (CEBR), King Saud University, Riyadh, Saudi Arabia