1. Department of Biology, Dire Dawa University, Dire Dawa, Ethiopia
2. Genetics and Plant Breeding Program, School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia
3. Plant Protection Program, School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia
4. United States Agriculture, Animal and Plant Health Inspection Service, Plant Protection and Quarantine, National Identification Service, Riverdale, MD, USA
5. Ethiopian Institute of Agricultural Research, Holetta Agricultural Research Center, Addis Ababa, Ethiopia