1. Key Laboratory of Sichuan Province for Fishes Conservation, Utilization in the Upper Reaches of the Yangtze River, Neijiang, Sichuan, PR China
2. College of Life Science, Neijiang Normal University, Neijiang, Sichuan, PR China
3. Fisheries Institute, Sichuan Academy of Agricultural Sciences, Chengdu, Sichuan, PR China
4. Research and Development Department, Sichuan Hengneng Fisheries Ltd, Neijiang, Sichuan, PR China