1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China
2. Yunnan Key Laboratory for Fungal Diversity and Green Development, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China
3. College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China
4. College of Life Sciences, Hunan Normal University, Changsha, Hunan 410000, China