1. Department of Biochemistry, Genetics and Microbiology, Forestry and Agricultural Biotechnology Institute (FABI), Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, 0028, South Africa
2. Protección Forestal, Centro de Investigación y Extensión Forestal Andino Patagónico, 9200 Esquel, Argentina
3. National Research Council of Argentina (CONICET), Buenos Aires, Argentina
4. Department of Plant and Soil Sciences, Forestry and Agricultural Biotechnology Institute (FABI), Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, 0028, South Africa