1. Department of Bioscience and Food Production Science, Interdisciplinary Graduate School of Science and Technology, Shinshu University, 8304, Minami-minowa, Nagano 399-4598, Japan
2. Sugadaira Research Station, Montane Science Center, University of Tsukuba, 1278-294, Sugadaira, Nagano 386-2204, Japan
3. Department of Bioscience and Biotechnology, Faculty of Agriculture, Shinshu University, 8304, Minami-minowa, Nagano 399-4598, Japan
4. Division of Terrestrial Ecosystem, Institute of Mountain Science, Shinshu University, 8304, Minami-minowa, Nagano 399-4598, Japan