1. Department of Medicine, Division of Infectious Diseases, Washington University School of Medicine, St. Louis, MO, USA
2. Edison Family Center for Genome Sciences & Systems Biology, Washington University School of Medicine, St. Louis, MO, USA
3. Department of Pathology & Immunology, Washington University School of Medicine, St. Louis, MO, USA
4. Division of Gastroenterology Addenbrooke’s Hospital and Department of Medicine, University of Cambridge, Cambridge, UK