1. Department of Applied Ecology, Faculty of Biology, St. Petersburg State University, St. Petersburg, Russia
2. Institute of Physico-Chemical and Biological Problems of Soil Science RAS, Pusсhino, Russia
3. V.L. Komarov Botany Institute RAS, Laboratory of Lichenology and Bryology, Saint-Petersburg, Russia
4. School of Environment and Sustainability, University of Saskatchewan, Saskatoon, Canada