1. School of Biology and Environmental Sciences, University of Mpumalanga, Nelspruit, South Africa
2. Aquatic Systems Research Group, Department of Geography and Environmental Science, University of Venda, Thohoyandou, South Africa
3. Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa
4. South African Institute for Aquatic Biodiversity, Makhanda, South Africa
5. Regional Integration Unit, Conservation Management, Skukuza, South Africa
6. Centre for Water Resources Research, University of KwaZulu–Natal, Pietermaritzburg, South Africa
7. Stellenbosch Institute for Advanced Study, Wallenberg Research Centre at Stellenbosch University, Stellenbosch, South Africa