1. a Institute of Microbiology , Leopold-Franzens University , Innsbruck, Technikerstraße 25, Innsbruck, A-6020, Austria
2. b INRA-CNRS , Station Recherches de Pathologie Comparée , St. Christol-les-Alès 30380, France
3. c School of Biological Sciences , University Wales Swansea , Singleton Park, Swansea, SA2 8PP, UK