1. State Key Laboratory of Grassland Agro-Ecosystems, College of Pastoral Agricultural Science and Technology, Lanzhou University, Lanzhou, China;
2. State Key Laboratory of Pharmaceutical Biotechnology, School of Life Science, Nanjing University, Nanjing, China;
3. BGI-Shenzhen, Shenzhen, China;
4. Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture, Lanzhou, China