1. Key Lab of Freshwater Biodiversity Conservation Ministry of Agriculture, Yangtze River Fisheries Research Institute, CAFS, Wuhan, China;
2. College of Animal Science, Guizhou University, Guiyang, China;
3. College of Animal Science and Technology, Northwest A&F University, Yangling, China;
4. Yellow River Fisheries Research Institute, Chinese Academy of Fishery Sciences, Xi’an, China