1. Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, PR China
2. Mécanismes Adaptatifs et Évolution (MECADEV), UMR 7179 CNRS–MNHN, Muséum national d’Histoire naturelle, CP50, 57 rue Cuvier, Paris, 75005, France